While pasta cooks, melt 3 tablespoons of the butter in a medium saucepan over medium. Whisk in flour, and cook, stirring constantly for 1 minute. Gradually pour in milk, whisking constantly until thoroughly combined. Bring to a boil, and cook, stirring often, until thickened, about 10 minutes. Remove from heat; stir in 6 ounces each of the blue cheese and the Cheddar, 1/2 teaspoon of the pepper, and remaining 1 teaspoon salt, stirring until cheese is melted. Pour cheese sauce over pasta; stir to coat. Transfer mac-and-cheese to a lightly greased 13- x 9-inch baking dish. Top evenly with remaining cheeses.
Place remaining 1 tablespoon butter in a small microwavable bowl; microwave until butter is melted, about 30 seconds. Stir in panko, lemon zest, and remaining 1/2 teaspoon black pepper.
Sprinkle breadcrumb topping over cheeses; bake in preheated oven until golden and bubbly, about 25 minutes. Sprinkle with parsley.
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